Copyright © 2018-2019, FFBO Teams All Rights Reserved.


The FFBO source code is licensed under the 3-Clause BSD License.


The FFBO documentation is licensed under the Creative Commons CC-BY 4.0 license.


The FFBO website is licensed under the Creative Commons CC-BY 4.0 license.


FFBO Teams

The FFBO is developed by the researchers across three continents. The members of each team are listed below.

  • The Columbia University Team
    • Wesley Bruning
    • Allison L. Fenichel
    • Aurel A. Lazar (PI)
    • Nikul H. Ukani
    • Chung-Heng Yeh
    • Yiyin Zhou
  • The NTHU Team
    • Ann-Shyn Chiang (PI)
    • Yu-Chi Huang
    • Chung-Chuang Lo (PI)
    • Cheng-Te Wang
  • The University of Sheffield Team
    • Daniel Coca (PI)
    • Dorian Florescu
    • Carlos Luna Ortiz
    • Paul Richmond
    • Adam Tomkins


The Open Science Prize Logo was obtained from and is licensed under the Creative Commons CC-BY 4.0 license.

The background image in the Under The Hood section is in public domain.


The development of FFBO has benefited enormously from the recent, focused efforts to produce connectome data in the fruit fly brain.

FlyCircuit Data

The FlyCircuit Database is provided by the NTHU Team. See [1] for detail.

[1] Ann-Shyn Chiang et al., Three-dimensional reconstruction of brain-wide wiring networks in Drosophila at single-cell resolution, Current Biology 2011, 21(1), pp.1-11

Hemibrain Data

Extracted from Hemibrain v1.0. See [2] for detail. Original data licensed under CC-BY.

[2] C. Shan Xu et al., A Connectome of the Adult Drosophila Central Brain. bioRxiv 2020.01.21.911859; doi:

Adult Drosophila Ventral Nerve Cord (VNC) Data

The data is retrieved from, January 11, 2020. See [3] for detail. The data is available under the Open Database License, with any rights in individual contents of the database are licensed under the Database Contents License.

[3] Jasper T. Maniates-Selvin et al., Reconstruction of motor control circuits in adult Drosophila using automated transmission electron microscopy. bioRxiv 2020.01.10.902478; doi:

Janelia Fly Medulla Data

The Fly Medulla data queriable in NeuroNLP is publicly available in this repository. The use of this data must follow the license:

This data cannot be used for biological discovery or biological publications before Fly EM publishes it. For other purposes (general bio data analysis algorithms, etc), this data is made available under the Open Data Commons Attribution License: A simple description of the license is provided here.

Fly Lamina Single Cartridge Data

We thank the authors of [2] to provide the data that the visualization of single cartridge neurons are based on.

[2] Marta Rivera-Alba et al., Wiring Economy and Volume Exclusion Determine Neuronal Placement in the Drosophila Brain, Current Biology 2011, 21(14), 2000–2005.